Negri performed extensive research in the fields of histology, hematology, aetiological agent of rabies was not a protozoan, but a filterable virus.
2.
Negri went on, however, to demonstrate in 1906 that the smallpox vaccine, then known as " vaccine virus ", or " variola vaccinae ", was also a filterable virus.
3.
By 1928 enough was known about viruses to enable the publication of " Filterable Viruses ", a collection of essays covering all known viruses edited by Thomas Milton Rivers ( 1888 1962 ).